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In the beginning, Medusa (メデューサ Medyuusa) co-ruled Angel Land with Palutena. Whereas Palutena fostered the growth of humanity, Medusa hated their kind, and took to drying up crops and petrifying people. Palutena, in a fit of anger, changed Medusa's form to a cyclopic, snake-haired monster and banished her to the Underworld. Despite this, or perhaps because of it, Medusa vowed one day to overthrow Palutena and retake her position in the Palace in the Sky. Over time, Medusa gained the allegiance of Underworld monsters and, once her forces were strong enough, led an attack against the very gates of the Palace. The ensuing war did not go well for Palutena; the Three Sacred Treasures were lost, Centurions were turned to stone, and Palutena herself was captured and imprisoned in the Palace in the Sky, unable to help as her beloved Angel Land swarmed with the forces of evil. As a last resort, she called out to Pit, the former leader of her personal bodyguards, now kept prisoner in the lowest dungeons of the Underworld. Responding to the call, and armed with a Bow manifested by the goddess' waning strength, Pit set out to destroy the invaders and defeat Medusa.
